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Young Brunette Girl With A Blue Feather
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Right: carotenoids (red) and melanins (dark) on belly/wings of Ramphocelus bresilius
The colors of feathers are produced by the presence of pigments, or by microscopic refractive structures, or by a combination of both.
Most feather pigments are melanins (brown and beige pheomelanins, black and grey eumelanins) and carotenoids (red, yellow, orange); other pigments occur only in certain taxa – the yellow to red psittacofulvins (found in some parrots) and the red turacin and green turacoverdin (porphyrin pigments found only in turacos). Structural coloration is involved in the production of blue colors, iridescence, most ultraviolet reflectance and in the enhancement of pigmentary colors; structural iridescence has been reported in fossil feathers dating back 40 million years. White feathers lack pigment and scatter light diffusely; albinism in birds is caused by defective pigment production, though structural coloration will not be affected (as can be seen e.g. in blue-and-white budgerigars).
For example, the blues and bright greens of many parrots are produced by constructive interference of light reflecting from different layers of the structures in feathers, in the case of green plumage in addition to the yellow pigments; the specific feather structure involved is sometimes called the Dyck texture. Melanin is often involved in the absorption of some of the light; in combination with yellow pigment it produces dull olive-greens.
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