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Brunette Girl Outside In The Deciduous Forest
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The converse of deciduous is evergreen, where green foliage is persistent year round. Plants that are intermediate, may be called semi-deciduous; they lose old foliage as new growth begins. Other plants are semi-evergreen and lose their leaves before the next growing season, but retain some during winter or during dry periods. Some trees, including a few species of oak have desiccated leaves that remain on the tree through winter; these dry persistent leaves are called marcescent leaves and are dropped in the spring as new growth begins.
Many deciduous plants flower during the period when they are leafless as this increases the effectiveness of pollination. The absence of leaves improves wind transmission of pollen for wind-pollinated plants, and increases the visibility of the flowers to insects in insect-pollinated plants. This strategy is not without risks, as the flowers can be damaged by frost or, in dry season regions, result in water stress on the plant. Nevertheless, there is much less branch and trunk breakage from glaze ice storms when leafless, and plants can reduce water loss due to the reduction in availability of liquid water during the cold winter days.
Leaf drop or abscission involves complex physiological signals and changes within plants. The process of photosynthesis steadily degrades the supply of chlorophylls in foliage; plants normally replenish chlorophylls during the summer months. When fall arrives and the days are shorter or when plants are drought stressed, deciduous trees decrease chlorophyll pigment production allowing other pigments present in the leaf to become apparent, resulting in non-green colored foliage. The brightest leaf colors are produced when days grow short and nights are cool but remain above freezing. These other pigments include carotenoids that are yellow, brown, and orange. Anthocyanin pigments produce reds and purple colors, though they are not always present in the leaves but are produced in the foliage in late summer when sugars are trapped in the leaves after the process of abscission begins. Parts of the world that have showy displays of bright autumn colors are limited to locations where days become short and nights are cool. In other parts of the world the leaves of deciduous trees simply fall off without turning the bright colors produced from the accumulation of anthocyanin pigments.
The beginning of leaf drop starts when an abscission layer is formed between the leaf petiole and the stem. This layer is formed in the spring during active new growth of the leaf, it consists of layers of cells that can separate from each other. The cells are sensitive to a plant hormone called auxin that is produced by the leaf and other parts of the plant. When the auxin coming from the leaf is produced at a rate consistent with that of the auxin from the body of the plant, the cells of the abscission layer remain connected; in autumn or when under stress the auxin flow from the leaf decreases or stops triggering cellular elongation within the abscission layer. The elongation of these cells break the connection between the different cell layers, allowing the leaf to break away from the plant, it also forms a layer that seals the break so the plant does not lose sap.
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