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When seeking to explain this change in the complexity and style of castles, antiquarians found their answer in the Crusades. It seemed that the Crusaders had learned much about fortification from their conflicts with the Saracens and exposure to Byzantine architecture. There were legends such as that of Lalys – an architect from Palestine who reputedly went to Wales after the Crusades and greatly enhanced the castles in the south of the country – and it was assumed that great architects such as James of Saint George originated in the East. However, in the mid-20th century this view was cast into doubt. Legends were discredited, and in the case of James of Saint George, it was proven that he came from Saint-Georges-d'Espéranche, in France. If the innovations in fortification had derived from the East, it would have been expected for their influence to be seen from 1100 onwards, immediately after the Christians were victorious in the First Crusade (1096–1099), rather than nearly 100 years later. Remains of Roman structures in Western Europe were still upstanding in many places, some of which had flanking round-towers and entrances between two flanking towers. The castle builders of Western Europe were aware of and influenced by Roman design; late Roman coastal forts on the English "Saxon Shore" were reused and in Spain the wall around the city of Ávila imitated Roman architecture when it was built in 1091. It has been argued – by historian Smail in Crusading warfare – that the case for the influence of Eastern fortification on the West has been overstated, and that Crusaders of the 12th century in fact learned very little about scientific design from Byzantine and Saracen defences. A well-sited castle that made use of natural defences and had strong ditches and walls had no need for a scientific design. An example of this approach is Kerak. Although there were no scientific elements to its design, it was almost impregnable, and in 1187 Saladin chose to lay siege to the castle and starve out its garrison rather than risk an assault.
After the First Crusade, Crusaders who did not return to their homes in Europe helped found the Crusader states of the principality of Antioch, the County of Edessa, the Kingdom of Jerusalem, and the County of Tripoli. The castles they founded to secure their acquisitions were designed mostly by Syrian master-masons. Their design was very similar to that of a Roman fort or Byzantine tetrapyrgia which were square in plan and had square towers at each corner that did not project much beyond the curtain wall. The keep of these Crusader castles would have had a square plan and generally be undecorated. While castles were used to hold a site and control movement of armies, in the Holy Land some key strategic positions were left unfortified. Castle architecture in the East became more complex around the late 12th and early 13th centuries after the stalemate of the Third Crusade (1189–1192). Both Christians and Muslims created fortifications, and the character of each was different. Saphadin, the 13th-century ruler of the Saracens, created structures with large rectangular towers that influenced Muslim architecture and were copied again and again, however they had little influence on Crusader castles.
In the early 13th century, Crusaders' castles were mostly built by Military Orders, such as the Knights Hospitaller, Knights Templar, and Knights of the Teutonic Order. They were responsible for the foundation of sites such as Krak des Chevaliers, Margat, and Belvoir. The forms of the castles varied not just between orders, but individually from castle to castle, although it was common for those founded in this period to have concentric defences. The concept, which originated in castles such as Krak des Chevaliers, was to remove the reliance on a central strongpoint and to emphasise the defence of the curtain walls. There would be more than one ring of defensive walls, one inside the other, with the inner ring rising above the outer so that its field of fire was not completely obscured. If assailants made it past the first line of defences into the outer enclosure, they would be caught in the killing ground between the inner and outer walls and have to assault the second wall to secure the fall of the castle. Concentric castles were widely copied across Europe, for instance when Edward I of England – who had himself been on Crusade – built castles in Wales in the late 13th century, four of the eight he founded were concentric. Not all the features of the Crusader castles from the 13th century were emulated in Europe; for instance, it was common in Crusader castles to have the main gate in the side of a tower and for there to be two turns in the passageway, lengthening the time it took for someone to reach the outer enclosure. It is rare for this bent entrance to be found in Europe.
One of the effects of the Livonian Crusade in the Baltic was the introduction of stone and brick fortifications. Although there were hundreds of wooden castles in Prussia and Livonia, the use of bricks and mortar was unknown in the region before the Crusaders. Until the 13th century and start of the 14th centuries, their design was heterogeneous, however this period saw the emergence of a standard plan in the region: a square plan, with four wings around a central courtyard. It was common for castles in the East to have arrowslits in the curtain wall at multiple levels; contemporary builders in Europe were wary of this as they believed it weakened the wall. Arrowslits did not compromise the wall's strength, but it was not until Edward I's programme of castle building that they were widely adopted in Europe. The Crusades also led to the introduction of machicolations into Western architecture. Until the 13th century, the tops of towers had been surrounded by wooden galleries, allowing defenders to drop objects on assailants below. Although machicolations performed the same purpose as the wooden galleries, they were probably an Eastern invention rather than an evolution of the wooden form. Machicolations were used in the East long before the arrival of the Crusaders, and perhaps as early as the first half of the 8th century in Syria. The greatest period of castle building in Spain was in the 11th to 13th centuries, and they were most commonly found in the disputed borders between Christian and Muslim lands. Conflict and interaction between the two groups led to an exchange of architectural ideas, and Spanish Christians adopted the use of detached towers. The Spanish Reconquista, driving the Muslims out of the Iberian Peninsula, was complete in 1492.